Cabezon (fish)
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The cabezon (''Scorpaenichthys marmoratus'') is a large species of
sculpin A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Scorpaeniformes.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology suggest transitions in station-holding demand a ...
native to the Pacific coast of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. Although the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
name translates literally as "scorpion fish", true scorpionfish (such as
lionfish ''Pterois'' is a genus of venomous marine fish, commonly known as lionfish, native to the Indo-Pacific. Also called firefish, turkeyfish, tastyfish, or butterfly-cod, it is characterized by conspicuous warning coloration with red, white, crea ...
) belong to the related family
Scorpaenidae The Scorpaenidae (also known as scorpionfish) are a family of mostly marine fish that includes many of the world's most venomous species. As their name suggests, scorpionfish have a type of "sting" in the form of sharp spines coated with venom ...
. The cabezon is the only known member of its genus.


Taxonomy

The cabezon was first formally described as ''Hemitripterus marmoratus'' in 1854 by the American
physician A physician (American English), medical practitioner (Commonwealth English), medical doctor, or simply doctor, is a health professional who practices medicine, which is concerned with promoting, maintaining or restoring health through th ...
and
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
William Orville Ayres William Orville Ayres (September 11, 1817 – April 30, 1887) was an American physician and ichthyologist. Born in Connecticut, he studied to become a doctor at Yale University School of Medicine. Life and career Ayers, the son of Jared and D ...
with its type locality given as California. Both Ayres and the French biologist
Charles Frédéric Girard Charles Frédéric Girard (8 March 1822 – 29 January 1895) was a French biologist specializing in ichthyology and herpetology. Born in Mulhouse, France, he studied at the College of Neuchâtel, Switzerland, as a student of Louis Agassiz. In ...
published the specific name ''marmoratus'' for this taxon in 1854, Ayres published his name on 8 September in ''The Pacific'', a San Francisco based journal in which the
California Academy of Sciences The California Academy of Sciences is a research institute and natural history museum in San Francisco, California, that is among the largest museums of natural history in the world, housing over 46 million specimens. The Academy began in 1853 ...
published its meeting repotrs, and the name was published once mor on 22 September in the ''
Proceedings of the California Academy of Sciences The California Academy of Sciences is a research institute and natural history museum in San Francisco, California, that is among the largest museums of natural history in the world, housing over 46 million specimens. The Academy began in 1853 ...
''. Girard's name was deemed to have been published on 6 October and authorship was confirmed in favour of Ayes in the
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the I ...
Opinion 1583 in 1990. Girard classified this species in the
monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
''Scorpaenichthys''. the cabezon is classified as belonging to the monotypic family Scorpaenichthyidae in the 5th edition of Fishes of the World. but subsquent authorities have placed the taxon within the
Jordaniidae Jordaniidae is a small family of marine ray-finned fishes belonging to the order Scorpaeniformes. These fishes are found in the eastern North Pacific Ocean. Taxonomy Jordaniidae was first proposed as a subfamily, Jordaniinae, of the family Cotti ...
. In either case the cabezon is regarded as one of the more basal members of the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Cottoidea A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Scorpaeniformes.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology suggest transitions in station-holding demand a ...
.


Description

The cabezon is a scaleless fish with a broad bony support extending from the eye across the cheek just under the skin. It has 11 spines on the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
. The cabezon also has a stout spine before the eye, an
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
of soft rays, and a fleshy flap on the middle of the
snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, rostrum, or proboscis. The wet furless surface around the nostrils of the nose of many mammals is c ...
. A pair of longer flaps are just behind the eyes. The mouth is broad with many small teeth. The coloring varies, but is generally mottled with browns, greens and reds. >90% of red fish are males, whereas >90% of green fish are females. The flesh is blue in color as are the internal organs. It reaches up to in length and in weight, while the largest ever Cabezon caught was in weight and longest being . As the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Can ...
-origin name implies, the fish has a very large head relative to its body.


Distribution and habitat

Cabezon are found in the northeast Pacific Ocean from
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
to
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
. They are found in a wide range of habitats at depths of , including rocky, muddy and sandy bottoms, and
kelp bed Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on Ea ...
s.


Fishing technique

Cabezon feed on
crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
,
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s, fish and fish eggs. Cabezon are taken as a
game fish Game fish, sport fish or quarry refer to popular fish pursued by recreational anglers, and can be freshwater or saltwater fish. Game fish can be eaten after being caught, or released after capture. Some game fish are also targeted commerciall ...
, but their roe is toxic to humans, because of the occurrence of a toxic
phospholipid Phospholipids, are a class of lipids whose molecule has a hydrophilic "head" containing a phosphate group and two hydrophobic "tails" derived from fatty acids, joined by an alcohol residue (usually a glycerol molecule). Marine phospholipids typ ...
( Dinogunellin). Cabezon inhabit the tops of rocky ledges as opposed to
rockfish Rockfish is a common term for several species of fish, referring to their tendency to hide among rocks. The name rockfish is used for many kinds of fish used for food. This common name belongs to several groups that are not closely related, and ca ...
and
lingcod The lingcod or ling cod (''Ophiodon elongatus''), also known as the buffalo cod or cultus cod, is a fish of the greenling family Hexagrammidae. It is the only extant member of the genus ''Ophiodon. ''A slightly larger, extinct species, '' Ophi ...
, which usually inhabit the sheer faces of these features.


References

*http://www.nmfs.noaa.gov/speciesid/fish_page/fish66a.html *International Game Fish Association "World Saltwater Records"
Occurrence of a toxic phospholipid in cabezon roe
{{Taxonbar, from=Q138228 Cottoidea Western North American coastal fauna Fish described in 1854